Sustained Flight


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How do insects stay in the air?

A series of experiments done by Michael Dickinson and his co-workers at the University of California Berkeley used fruit flies (Diptera: Drosophila) and a large scale Robofly wing in oil to answer this question.

By watching the live flies researchers divided each stroke of an Insect's wings into two translational phases and two rotational phases.  Translational phases, one up and one down, are where the insect would push their wings through the air generating force.  Rotational phases, pronation and supination, involve rotational movement of the wings in order to ready the insect's wing for the next translational phase.

The same UC Berkeley researchers then used the large-scale Robofly wing in oil to determine the estimated magnitude and direction of the forces on the wing (red arrows in the image below) as well as the forces created by the insect (blue arrows in the image below).  Oil was used in order to simulate the effects of air on the fruit fly wing relative to its actual size.
 
Image showing the
      forces on and created by a fruit fly wing
Image showing the forces on and created by a fruit fly wing
http://www.scientificamerican.com/media/inline/00029B82-5453-1C75-9B81809EC588EF21_arch3.gif

The result was the discovery of three ways in which insects are able to sustain flight through lift.
Delayed Stall, Rotational Circulation, and Wake Capture

More details
and video (off site links)

Another series of experiments preformed by Dr. John Young of the University of New South Wales, Australia in collaboration with researchers from Oxford University placed locusts (Orthoptera) in a wind tunnel and filmed them with high-speed digital cameras.  Locusts were chosen because they have incredible conservation of energy during flight as evidenced by their ability to fly large distances with extremely limited energy reserves.

"Until very recently it hasn't been possible to measure the actual shape of an insect's wings in flight – partly because their wings flap so fast, and partly because their shape is so complicated." - Dr. John Young

The results of their research showed how the shape of a locust's wing changed during flight.  These changes are extremely important in counteracting the Force of gravity (F=m*g) on the insect by allowing it to generate maximum lift. 

Smoke visualization of locust in wind
        tunnel
Smoke visualization of locust in wind tunnel
http://www.sciencedaily.com/releases/2009/09/090917144125.htm

Practical applications of this research included creating a computer model which allows researchers to see a replicated airflow and the resulting thrust generated by the locust, and to disprove the 'bumblebee paradox' which states that insects defy the laws of aerodynamics.  This computer model will also be useful in developing insect-like micro-air robots such as Robofly.

"The message for engineers working to build insect-like micro-air vehicles is that the high lift of insect wings may be relatively easy to achieve, but that if the aim is to achieve efficiency of the sort that enables inter-continental flight in locusts, then the details of deforming wing design are critical." - Dr. John Young